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Clark University    
  Feb 20, 2018
2017-2018 Academic Catalog

Biology, PhD

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Graduate Program

The department offers course work and research opportunities leading to the Doctor of Philosophy in Biology. The department has two foci for graduate emphasis: molecular and cell biology or ecology and evolution.

Students applying for admission must demonstrate strong preparation in biology and satisfactory scores on the Graduate Record Examination (and TOEFL for International Students). Tuition scholarships and research and teaching assistantships are available to qualified students. Further information, including course requirements, application procedures, and policies can be obtained from the Biology Web site at

Biology Faculty


Nathan Ahlgren, Ph.D.

Philip J. Bergmann, Ph.D.

Robert Drewell, Ph.D.

Susan Foster, Ph.D.

John Gibbons, Ph.D.

David Hibbett, Ph.D.

Denis Larochelle, Ph.D.

Néva Meyer, Ph.D.

Deborah Robertson, Ph.D.

Justin Thackeray, Ph.D.


Halina Brown, Ph.D.
Frederick Greenaway, Ph.D.
Dominik Kulakowski, Ph.D.
Noel Lazo, Ph.D.
John Rogan, Ph.D.
Donald Spratt, Ph.D.
David Thurlow, Ph.D.
Christopher A. Williams, Ph.D.


John Baker, Ph.D.


John Brink, Ph.D.

Linda Kennedy, Ph.D.

Thomas Leonard, Ph.D.

Todd Livdahl, Ph.D.

Timothy Lyerla, Ph.D.

John Reynolds, Ph.D.

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