2012-2013 Academic Catalog 
    Mar 23, 2018  
2012-2013 Academic Catalog [ARCHIVED CATALOG]

Biology, PhD


Graduate Program

The department offers course work and research opportunities leading to the Doctor of Philosophy in Biology. The department has two foci for graduate emphasis: molecular and cell biology or ecology and evolution.

Students applying for admission must demonstrate adequate preparation in the basic sciences, an overall undergraduate record of B– or better, and satisfactory scores on the Graduate Record Examination (and TOEFL for International Students). Tuition scholarships and research and teaching assistantships are available to qualified students. Further information can be obtained from the Biology Web site at http://www.clarku.edu/departments/biology/.

Course Requirements

Each student’s Advisory Committee will determine which courses must be taken, based on the student’s research interests and prior training. These requirements, as well as the residency requirement, will be met with a grade of B- or better. Grades lower than B- will not be counted towards the degree, and students who accumulate more than two grades lower than B- will not be allowed to continue in the program. In addition to formal courses stipulated by the committee, graduate students are required to enroll in the Graduate Research Seminar (BIOL 350) and attend all departmental seminars.

Biology Faculty


Philip J. Bergmann, Ph.D.
Susan Foster, Ph.D.
David Hibbett, Ph.D.
Denis Larochelle, Ph.D.
Todd Livdahl, Ph.D.
Néva Meyer , Ph.D.
Deborah Robertson, Ph.D.
Justin Thackeray, Ph.D.
Heather Wiatrowski, Ph.D.


Halina Brown, Ph.D.
Frederick Greenaway, Ph.D.
Dominik Kulakowski, Ph.D.
Noel Lazo, Ph.D.
John Rogan, Ph.D.
Joseph Tang, Ph.D.
David Thurlow, Ph.D.
Christopher A. Williams, Ph.D.


John Baker, Ph.D.
Linda Kennedy, Ph.D.


John Brink, Ph.D.
Joseph Curtis, Ph.D.
H. William Johansen, Ph.D.
Thomas Leonard, Ph.D.
Timothy Lyerla, Ph.D.
John Reynolds, Ph.D.